| Phrynobatrachus | |
|---|---|
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| A male and a female of Phrynobatrachus bibita | |
| Scientific classification  | |
| Domain: | Eukaryota | 
| Kingdom: | Animalia | 
| Phylum: | Chordata | 
| Class: | Amphibia | 
| Order: | Anura | 
| Clade: | Ranoidea | 
| Family: | Phrynobatrachidae Laurent, 1941  | 
| Genus: | Phrynobatrachus Günther, 1862  | 
| Species | |
| 
 See text  | |
| Synonyms | |
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Phrynobatrachus is a genus of Sub-Saharan frogs that form the monogeneric family Phrynobatrachidae. Their common name is puddle frogs, dwarf puddle frogs, African puddle frogs, or African river frogs.[1][2][3][4] The common name, puddle frog, refers to the fact that many species breed in temporary waterbodies such as puddles.[5]
Phrynobatrachus are among the most common amphibians in Africa. They are typically small (mostly less than 30 mm (1.2 in)[5]), fast-moving frogs. They occupy a variety of habitats from dry savannas to rainforests. Most species deposit many small eggs as a surface clutch in standing or slowly moving water and have exotrophic tadpoles.[4]
Taxonomy
Phrynobatrachidae has earlier been considered as a subfamily of Ranidae, but its recognition as a family is now well-established.[1][2][3][4] It is probably most closely related to Petropedetidae and Pyxicephalidae[1] or Ptychadenidae.[4]
This large genus may be further divided into three major clades. These clades could be treated as different genera, but this arrangement is not yet in use.[1]
Species
There are currently 96 species in this genus:[6]
- Phrynobatrachus acridoides (Cope, 1867)
 - Phrynobatrachus acutirostris Nieden, 1912
 - Phrynobatrachus afiabirago Ofori-Boateng, Leaché, Obeng-Kankam, Kouamé, Hillers & Rödel, 2018
 - Phrynobatrachus africanus (Hallowell, 1858)
 - Phrynobatrachus albifer (Ahl, 1924)
 - Phrynobatrachus albomarginatus De Witte, 1933
 - Phrynobatrachus alleni Parker, 1936
 - Phrynobatrachus ambanguluensis Greenwood, Loader, Lawson, Greenbaum, and Zimkus, 2020
 - Phrynobatrachus amieti Nečas, Dolinay, Zimkus, and Gvoždík, 2021
 - Phrynobatrachus annulatus Perret, 1966
 - Phrynobatrachus anotis Schmidt and Inger, 1959
 - Phrynobatrachus arcanus Gvoždík, Nečas, Dolinay, Zimkus, Schmitz, and Fokam, 2020
 - Phrynobatrachus asper Laurent, 1951
 - Phrynobatrachus auritus Boulenger, 1900
 - Phrynobatrachus batesii (Boulenger, 1906)
 - Phrynobatrachus bequaerti (Barbour and Loveridge, 1929)
 - Phrynobatrachus bibita Goutte, Reyes-Velasco, and Boissinot, 2019
 - Phrynobatrachus breviceps Pickersgill, 2007
 - Phrynobatrachus brevipalmatus (Ahl, 1925)
 - Phrynobatrachus brongersmai Parker, 1936
 - Phrynobatrachus bullans Crutsinger, Pickersgill, Channing, and Moyer, 2004
 - Phrynobatrachus calcaratus (Peters, 1863)
 - Phrynobatrachus chukuchuku Zimkus, 2009
 - Phrynobatrachus congicus (Ahl, 1925)
 - Phrynobatrachus cornutus (Boulenger, 1906)
 - Phrynobatrachus cricogaster Perret, 1957
 - Phrynobatrachus cryptotis Schmidt and Inger, 1959
 - Phrynobatrachus dalcqi Laurent, 1952
 - Phrynobatrachus danko Blackburn, 2010
 - Phrynobatrachus dendrobates (Boulenger, 1919)
 - Phrynobatrachus discogularis Pickersgill, Zimkus, and Raw, 2017
 - Phrynobatrachus dispar (Peters, 1870)
 - Phrynobatrachus elberti (Ahl, 1925)
 - Phrynobatrachus francisci Boulenger, 1912
 - Phrynobatrachus fraterculus (Chabanaud, 1921)
 - Phrynobatrachus gastoni Barbour and Loveridge, 1928
 - Phrynobatrachus ghanensis Schiøtz, 1964
 - Phrynobatrachus giorgii De Witte, 1921
 - Phrynobatrachus graueri (Nieden, 1911)
 - Phrynobatrachus guineensis Guibé and Lamotte, 1962
 - Phrynobatrachus gutturosus (Chabanaud, 1921)
 - Phrynobatrachus hieroglyphicus Rödel, Ohler, and Hillers, 2010
 - Phrynobatrachus horsti Rödel, Burger, Zassi-Boufou, Emmrich, Penner, and Barej, 2015
 - Phrynobatrachus hylaios Perret, 1959
 - Phrynobatrachus inexpectatus Largen, 2001
 - Phrynobatrachus intermedius Rödel, Boateng, Penner, and Hillers, 2009
 - Phrynobatrachus irangi Drewes and Perret, 2000
 - Phrynobatrachus jimzimkusi Zimkus, Gvoždík, and Gonwouo, 2013
 - Phrynobatrachus kakamikro Schick, Zimkus, Channing, Köhler, and Lötters, 2010
 - Phrynobatrachus keniensis Barbour and Loveridge, 1928
 - Phrynobatrachus kinangopensis Angel, 1924
 - Phrynobatrachus krefftii Boulenger, 1909
 - Phrynobatrachus latifrons Ahl, 1924
 - Phrynobatrachus leveleve Uyeda, Drewes, and Zimkus, 2007
 - Phrynobatrachus liberiensis Barbour and Loveridge, 1927
 - Phrynobatrachus mababiensis FitzSimons, 1932
 - Phrynobatrachus maculiventris Guibé and Lamotte, 1958
 - Phrynobatrachus manengoubensis (Angel, 1940)
 - Phrynobatrachus mayokoensis Rödel, Burger, Zassi-Boulou, Emmrich, Penner, and Barej, 2015
 - Phrynobatrachus mbabo Gvoždík, Nečas, Dolinay, Zimkus, Schmitz, and Fokam, 2020
 - Phrynobatrachus minutus (Boulenger, 1895)
 - Phrynobatrachus nanus (Ahl, 1925)
 - Phrynobatrachus natalensis (Smith, 1849)
 - Phrynobatrachus njiomock Zimkus and Gvoždík, 2013
 - Phrynobatrachus ogoensis (Boulenger, 1906)
 - Phrynobatrachus pakenhami Loveridge, 1941
 - Phrynobatrachus pallidus Pickersgill, 2007
 - Phrynobatrachus parkeri De Witte, 1933
 - Phrynobatrachus parvulus (Boulenger, 1905)
 - Phrynobatrachus perpalmatus Boulenger, 1898
 - Phrynobatrachus petropedetoides Ahl, 1924
 - Phrynobatrachus phyllophilus Rödel and Ernst, 2002
 - Phrynobatrachus pintoi Hillers, Zimkus, and Rödel, 2008
 - Phrynobatrachus plicatus (Günther, 1858)
 - Phrynobatrachus pygmaeus (Ahl, 1925)
 - Phrynobatrachus rainerguentheri Rödel, Onadeko, Barej, and Sandberger, 2012
 - Phrynobatrachus rouxi (Nieden, 1912)
 - Phrynobatrachus rungwensis (Loveridge, 1932)
 - Phrynobatrachus ruthbeateae Rödel, Doherty-Bone, Kouete, Janzen, Garrett, Browne, Gonwouo, Barej, and Sandberger, 2012
 - Phrynobatrachus sandersoni (Parker, 1935)
 - Phrynobatrachus scapularis (De Witte, 1933)
 - Phrynobatrachus scheffleri (Nieden, 1911)
 - Phrynobatrachus schioetzi Blackburn and Rödel, 2011
 - Phrynobatrachus steindachneri Nieden, 1910
 - Phrynobatrachus sternfeldi (Ahl, 1924)
 - Phrynobatrachus stewartae Poynton and Broadley, 1985
 - Phrynobatrachus sulfureogularis Laurent, 1951
 - Phrynobatrachus taiensis Perret, 1988
 - Phrynobatrachus tanoeensis Kpan, Kouamé, Barej, Adeba, Emmrich, Boateng, and Rödel, 2018
 - Phrynobatrachus tokba (Chabanaud, 1921)
 - Phrynobatrachus ukingensis (Loveridge, 1932)
 - Phrynobatrachus ungujae Pickersgill, 2007
 - Phrynobatrachus uzungwensis Grandison and Howell, 1983
 - Phrynobatrachus versicolor Ahl, 1924
 - Phrynobatrachus villiersi Guibé, 1959
 - Phrynobatrachus werneri (Nieden, 1910)
 
References
- 1 2 3 4 Frost, Darrel R. (2014). "Phrynobatrachidae Laurent, 1941". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 10 May 2014.
 - 1 2 "Phrynobatrachidae". AmphibiaWeb: Information on amphibian biology and conservation. [web application]. Berkeley, California: AmphibiaWeb. 2014. Retrieved 10 May 2014.
 - 1 2 Blackburn, D. C.; Wake, D. B. (2011). "Class Amphibia Gray, 1825. In: Zhang, Z.-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness" (PDF). Zootaxa. 3148: 39–55. doi:10.11646/zootaxa.3148.1.8.
 - 1 2 3 4 Vitt, Laurie J.; Caldwell, Janalee P. (2014). Herpetology: An Introductory Biology of Amphibians and Reptiles (4th ed.). Academic Press. p. 507.
 - 1 2 Zimkus, B. "Phrynobatrachus Günther, 1862". African Amphibians Lifedesk. Archived from the original on 6 October 2011. Retrieved 10 May 2014.
 - ↑ Frost, Darrel R. (2022). "Phrynobatrachus Günther, 1862". Amphibian Species of the World: an Online Reference. Version 6.0. American Museum of Natural History. Retrieved 28 July 2022.
 
External links
 Media related to Phrynobatrachus at Wikimedia Commons
